Classification of Caucasian oak (Quercus macranthera) forests from Dagestan (Eastern Caucasus)
UDC 581.526.425:581.559(470.67)
Abstract
A classification of Caucasian oak (Quercus macranthera) forests from Dagestan (Eastern Caucasus) using the Braun-Blanquet method has been developed. Three new associations were described: Ranunculo buhsei–Quercetum macrantherae ass. nov., Betonico macrantherae–Quercetum macrantherae ass. nov. and Calamagrostio arundnaceae–Quercetum macrantherae ass. nov. These communities are characterized by the leading role of sub-Mediterranean thermophilous species and Euro-Siberian meadow-steppe meso-xerophytes which allowed them to be assigned to the class Quercetea pubescentis Doing-Kraft ex Scamoni et Passarge 1959. The results of the study of ecological and plant-geographical properties of described communities revealed their unique feature – the presence of a large group of endemic subalpine Caucasian species. The combination of these ecologically contrasting species groups indicates moderately dry and continental climate of the oak (Q. macranthera) forests range in Dagestan and occurring them in the buffer zone between forest, steppe and alpine belts at altitudes of 1400–2100 m. Similar important ecological and floristic features are not typical for other forests of the class Quercetea pubescentis in Europe and Asia Minor. This made it possible to justify a new order Astrantio maximae–Quercetalia macrantherae ord. nov. containing a single alliance Astrantio maximae–Quercion macrantherae all. nov. Diagnostic species of the order and alliance are: Quercus macranthera, Astrantia maxima, Alchemilla sericata, Anthemis melanoloma, Betonica macrantha, Bupleurum polyphyllum, Calamagrostis arundinacea, Carum meifolium, Chaerophyllum aureum, Cephalaria gigantea, Centaurea salicifolia, Cicerbita racemosa, Erysimum armeniacum, Geranium sylvaticum, Heracleum asperum, Iris colchica, Lathyrus cyaneus, Lilium monadelphum, Medicago glutinosa, Primula macrocalyx, Viburnum lantana, Poa longifolia, Polygonatum verticillatum, Ranunculus buhsei, Rubus saxatilis, Primula ruprechtii, Psephellus dealbatus, P. daghestanicus, Pyrethrum leptophyllum, Pedicularis chroorrhyncha, Sedum oppositifolium, Trifolium canescens, Vicia grossheimii, Vincetoxicum funebre.
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Akman Y., Barbero M., Quezel P. 1979. Contribution to the study of forest vegetation in Mediterranean Anatolia. Phytocoenologia 5: 1–79. [In French]. DOI: 10.1127/phyto/5/1978/1
Cansaran A., Kaya Ö. F., Ertekin A. S., Ketenoğlu O. A. 2010. A phytosociological study on Karaömer Mountain of North Anatolia (Amasya, Turkey). Acta Bot. Gallica 157(1): 65–88. DOI: 10.1080/12538078.2010.10516190
Coban S., Willner W. 2019. Numerical classification of the forest vegetation in the Western Euxine Region of Turkey. Phytocoenologia 49, 1: 71–106. DOI: 10.1127/phyto/2018/0274
Czerepanov S. K. 1995. Vascular plants of Russia and adjacent states (the former USSR). Cambridge: Cambridge University Press. 516 pp.
Долуханов А. Г. Типологический очерк горных лесов из грузинского и восточного дуба // Труды Тбилисского ботанического института, 1955. Т. 17. С. 73–160.
Ermakov N. B., Plugatar Yu. V., Leiba V. D. 2020. Endemic Quercus pontica C. Koch. communities from the Colchic Province and new syntaxonomical concept for the Caucasian subalpine krummholz vegetation. Botanica Pacifica. A journal of plant science and conservation 9, 2: 1–9.
Gholizadeh H., Naqinezhad A., Chytrý M. 2019. Hyrcanian forest vegetation database. Phytocoenologia 49: 209–210. DOI: 10.1127/phyto/2018/0315
Gholizadeh H., Naqinezhad A., Chytrý M. 2020. Classification of the Hyrcanian forest vegetation, northern Iran. Applied Vegetation Science 23: 107–126. DOI: 10.1111/avsc.12469
Гулисашвили В. З., Махатадзе Л. Б., Прилипко Л. И. Растительность Кавказа. М.: Наука, 1985. 233 с.
Hennekens S. M., Schaminée J. H. J. 2001: TURBOVEG, a comprehensive data base management system for vegetation data. Journal of Vegetation Science 12: 589–591. DOI: 10.2307/3237010
Ignatov M. S., Afonina O. M., Ignatova E. A., Abolina A., Akatova T. V., Baisheva E. Z., et al. 2006. Check-list of mosses of East Europe and North Asia. Arctoa 15: 1–130. DOI: 10.15298/arctoa.15.01
Karaer F., Kilinc M., Kutbay H. G. 1999. The Woody Vegetation of the Kelkit Valley. Turkish Journal of Botany 23: 319–344.
Kaya Ö. F., Cansaran A., Yildinm C. 2010. A syntaxonomical investigation of forest and pseudomaquis on transitional area in the central Black Sea region (Amasya, Turkey). Acta Bot. Gallica 157(3): 469–482. DOI: 10.1080/12538078.2010.10516223
Kenar N. 2017. Phytosociological investigations of steppe and steppe forest vegetation in the south-east part of Central Anatolia of Turkey. Journal of the Faculty of Forestry Istanbul University 67(2): 203–219. DOI: 10.17099/jffiu.322422
Korkmaz H., Engin A., Kutbay H. G., Yalcin E. 2011. A syntaxonomical study on the scrub, forest, and steppe vegetation of the Kızılırmak valley. Turkish Journal of Botany 35: 121–165. DOI: 10.3906/bot-0908-152
Меницкий Ю. Л. Дубы Азии. Л.: Наука, 1984. 316 с.
Özen F., Kilinc M. 2002. The Flora and Vegetation of Kunduz Forests (Vezirköprü / Samsun). Turkish Journal of Botany 26: 371–393.
Passarge H. 1981. Carpineta in kartalinischen Kaukasus. Phytocoenologia 9: 533–545.
Quézel P., Barbero M., Akman Y. 1980. Contribution ál’étude de la végétation forestière d’Anatolie septentrionale. [Contribution to the study of forest vegetation of Northern Anatolia]. Phytocoenologia 8(3–4): 365–519. DOI: 10.1127/phyto/8/1980/365
Theurillat J.-P., Willner W., Fernández-González F., Bültmann H., Čarni A., Gigante G., Mucina L., Weber H. 2020. International Code of Phytosociological Nomenclature. 4th edition. Applied Vegetation Science. DOI: 10.1111/avsc.12491
Tichý L. 2002. JUICE, software for vegetation classification. Journal of Vegetation Science 13: 451–453. DOI: 10.1111/j.1654–1103.2002.tb02069.x
Ture C., Tokur S., Ketenoglu O. 2005. Contributions to the syntaxonomy and ecology of the forest and shrub vegetation in Bithynia, Northwestern Anatolia, Turkey. Phyton 45(1): 81–115.
Westhoff V., van der Maarel E. 1973. The Braun-Blanquet Approach. In: R. H. Whittaker (ed.). Ordination and classification of communities. Dordrecht: Dr. W. Junk. Pp. 617–626.
Yildirim C., Erkan Yalcin E., Cansaran A., Korkmaz H. 2019. Syntaxonomic analysis of forests, shrubs, and steppes of Tavşan Mountain (Amasya, Turkey). Turkish Journal of Botany 43: 409–419. DOI: 10.3906/bot-1809-18
Zohary M. 1963. On the geobotanical structure of Iran. Jerusalem: Weizmann Science Press of Israel. 113 pp.
Zohary M. 1973. Geobotanical foundations of the Middle East. Vol. 1–2. Stuttgart: Gustav Fischer Verlag. 739 pр.
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