Korolyuk, E. A., M. N. Lomonosova, and T. E. Aleshina. “Karyotypes of Two Endemic Species of Asterothamnus Novopokr. (Asteraceae) from South Siberia (Tuva)”. Turczaninowia, Vol. 19, no. 3, Sept. 2016, pp. 115-9, https://turczaninowia.asu.ru/article/view/1513.